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MU.LAB has 2 different kinds of focus:
The GUI Focus defines which GUI component will process the keys you press on your computer keyboard. More info here.
The MIDI Input Focus defines which plug-in will process the MIDI input events. MU.LAB indicates the MIDI input focus by highlighting the plug-in.
By default, MU.LAB syncs these two focusses.
But there are situations where it is better to not have them synced: Imagine you're making music with someone and you are editing on the computer and the other person is playing a MIDI keyboard. Now whenever you click on a plug-in on the screen the MIDI input focus is shifted to that plug-in and so the sound on the person's keyboard will change. That's not comfortable. So that's why you can tell MU.LAB to "Use Manual MIDI Input Focus". Then whenever you do want to change the MIDI input focus, choose "Play From MIDI Input" from a module's context menu 
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